These small twelfth stater (Obol) from Miletos Ionia are one of the cities most familiar symbols. The Ionian region, located in what is now Turkey, was a narrow strip along the coast from Phocaea to the mouth of Hermus River, now the Gediz, to Miletos, Ionia at the mouth of the Maender River and included the two islands of Chios and Samos. The cities within this informal region were of great importance in the conflicts between the Greeks and the Persian Empire. The Greeks claimed that the Ionian region was settled by peoples from the other side of the Aegean Sea. According to the Greeks the Ionic settlers of the region were led by Neleus and Androelus, the sons of the last King of Athens, Codrus. The defeat of the Persian Empire at the battle of Granicus by Alexander the Great brought the region under the control of the Macedonians (Greeks). Most of the cities of the region submitted to Alexander the Greats rule and enjoyed great prosperity during this (Hellenistic) period, except for Miletos Ionia, which refused to pay homage to Alexander the Great and was eventually leveled in 334 BC and never returned to its pervious glory.